Invasive species are non-native organisms establishing self-sustaining populations and negatively impacting native biodiversity or ecosystem function. Establishment depends on propagule pressure, species' invasiveness traits, and recipient community resistance. Invasive species outcompete natives, alter disturbance regimes, or disrupt trophic networks.
Not every non-native species becomes invasive. From your understanding of species interactions and community ecology, you know that communities resist newcomers through competition, predation, and resource limitation. An invasive species is one that overcomes these barriers, establishes a self-sustaining population, and causes measurable harm to native biodiversity or ecosystem processes. The critical distinction is between a species that merely survives in a new location and one that spreads aggressively and reshapes the community around it.
Propagule pressure — the number of individuals introduced and the frequency of introduction events — is the strongest predictor of whether a non-native species establishes. A single seed blown across an ocean rarely founds an invasion; a thousand seeds dumped repeatedly in ballast water often do. Once enough individuals arrive, the probability that at least some survive environmental filtering, find mates, and reproduce rises sharply. This is why shipping ports, garden centers, and aquaculture facilities are invasion hotspots: they deliver high propagule pressure repeatedly.
Successful invaders often share a suite of invasiveness traits: rapid reproduction, broad environmental tolerance, high dispersal ability, and phenotypic plasticity. But the invader's traits alone do not determine success — the invasibility of the recipient community matters equally. Communities with low species diversity, disturbed habitats, or available empty niches offer less biotic resistance. This is why islands, which you know from biodiversity and conservation studies tend to have fewer native competitors and predators, are disproportionately vulnerable to invasions.
Once established, invasive species disrupt native communities through several mechanisms. Competitive displacement occurs when an invader monopolizes resources more efficiently than natives — the zebra mussel filtering plankton faster than native mussels, for example. Trophic disruption occurs when an invader becomes a new predator (like the brown tree snake eliminating Guam's native birds) or removes a key prey species. Some invaders act as ecosystem engineers, fundamentally altering the physical environment: invasive grasses in Hawaiian forests increase fire frequency, converting fire-intolerant native forest to fire-adapted grassland in a positive feedback loop. These cascading effects mean that a single invader can restructure entire food webs and nutrient cycles, connecting back to the trophic network concepts you already understand.
The severity of invasion impacts depends on context. Not all non-native species cause harm, and some integrate into communities without measurable damage. The challenge for conservation biology is distinguishing harmful invaders from benign introductions early enough to act, since eradication becomes exponentially harder as populations grow and spread. Prevention — controlling propagule pressure through trade regulation and biosecurity — remains far more cost-effective than removal after establishment.